Full CV is available here.

In situ

Willoughby, E. A., & Lee, James J. (Under review). Parsing information flow in speeded cognitive tasks.

Anderson, E. L., Saunders, G. R. B., Willoughby, E. A., Iacono, W. G., McGue, M. (Under review). Uncovering the genetic and environmental mediators of parent-offspring transmission of educational attainment: An adoption study. Under review at Journal of Personality.

Willoughby, E. A. & Lee, J. J. (In press). Predicting cognitive ability differences from brain imaging and genetics data. In: The Cambridge handbook of intelligence and neuroscience.

Willoughby, E. A. (In situ). Drawing and painting dinosaurs. Marlborough, UK: The Crowood Press. Projected release in 2021.


McGue, M., Willoughby, E. A., Rustichini, A., Johnson, W., Iacono, W., & Lee, J. J. (2020). The contribution of cognitive and non-cognitive skills to intergenerational social mobility. Psychological Science, 31(7), 835–847. DOI: 10.1177/0956797620924677

Gulsuner, S., Stein, D. J., Susser, E. S., Sibeko, G., Pretorius, A., Walsh, T., Majara, L., Mndini, M. M., Mqulwana, S. G., Ntola, O. A., Casadei, S., Ngqengelele, L. L., Korchina, V., van der Merwe, C., Malan, M., Fader, K. M., Feng, M., Willoughby, E. A. . . . & McClellan, J. M. (2020). Genetics of schizophrenia in the South African Xhosa. Science, 367(6477), 569–573. DOI: 10.1126/science.aay8833

Willoughby, E. A., McGue, M., Iacono, W., Rustichini, A., & Lee, J. J. (2020). The role of parental genotype in predicting offspring years of education: Evidence for genetic nurture. Molecular Psychiatry, online first. DOI: 10.1038/s41380-019-0494-1


Willoughby, E. A., Love, A. C., McGue, M., Iacono, W. G., Quigley, J., & Lee, J. J. (2019). Free will, determinism, and intuitive judgments about the heritability of behavior. Behavior Genetics, 49(2), 136–153. DOI: 10.1007/s10519-018-9931-1


Willoughby, E. A. & Boutwell, B. B. (2018). Importance of intelligence and emotional intelligence for physicians. Journal of the American Medical Association, 320(2), 205. DOI: 10.1001/jama.2018.6278

Lee, J. J., Wedow, R., Okbay, A., Kong, E., Maghzian, O., Zacher, M., Nguyen-Viet, T. A., Bowers, P., Sidorenko, J., Linner, R. K., Fontana, M. A., Kundu, T., Lee, C., Li, H., Li, R., Royer, R., Timshel, P. N., Walters, R. K., Willoughby, E. A. . . . Cesarini, D. (2018). Gene discovery and polygenic prediction from a 1.1-million-person GWAS of educational attainment. Nature Genetics, 50(8), 1112–1121. DOI: 10.1038/s41588-018-0147-3


Kane, J., Willoughby, E. & Keesey, M. (2016). God’s word or human reason? Portland, Oregon: Inkwater Press. ISBN: 978-1629013725. (Author & lead illustrator)